Gulping down oysters has long been a favorite New Year's Eve ritual for the French, but as winters get warmer and summers get drier many growers worry there will soon be fewer of the prized mollusks to go around.

"Twenty years ago, we'd be shivering in the warehouse while preparing the holiday orders. Today it's 15 degrees Celsius (60 degrees Fahrenheit)," says Brittany oysterman Mathieu Le Moal, his sleeves rolled up in front of a tractor carrying dozens of bulging oyster sacks.

"We don't have seasons anymore - but oysters need all four," Le Moal adds. "They need the winter, it's when they can rest, use less energy."

Inside a wooden hanger redolent of salt and the sea, around a dozen of his workers are sorting, weighing and packing oysters into crates in the Brittany port of Cancale.

Le Moal and other farmers along this stretch of France's Emerald Coast say the long drought which struck swathes of the country this summer took a heavy toll, leading to smaller harvests, and smaller shellfish.

Without summer rains that wash crucial minerals into the oyster beds, "there's no plankton, the main food for oysters, so they don't grow," explained fellow oysterman Bertrand Racinne, weaving his way between baskets and stacked crates.

"In the end, we have oysters but not enough of the big ones," said Racinne, who like most growers sells more than half his yearly production in December. Cold weather normally encourages a needed rest for oysters to mature, said Yoann Thomas of France's IRD research institute.

But this winter has so far been unusually warm and, paradoxically, too rainy.

Rains may bring minerals that favor plankton growth - but they also mean the mollusks spend too much energy eating. This year's harvest are likely to start the spring "fragile and vulnerable," warned Racinne.

"We've found that periods of extreme mortality (more than 25 percent of oysters) come several months after mild and rainy winters," Thomas said.

Germs thrive

"Ten grams fewer for each one, that makes a difference in sales," said Philippe Le Gal, president of the CNC national shellfish producers' association.

In 2017 the roughly 4,500 oyster growers in France sold 100,000 tons, at an average price of 5,000 euros ($5,700) per ton.

"Oyster farmers will see volumes down by 20 to 30 percent this year," Le Gal said. "Climate warming is starting to have an impact."

Warmer water temperatures are also a risk because they facilitate the spread of viruses that are especially harmful to oyster larvae, or spat, and young oysters.

Scientists point in particular to a Herpes virus, OsHV-1, which has been present in French oyster waters since 1991 but has become more aggressive, recently, for reasons still unknown.

Since 2008, up to 75 percent of young oysters have been lost in some years, said Fabrice Pernet at the Ifremer ocean research institute in Brest.

"Oyster farmers had found a solution by putting 10 times the amount of spat in the water in autumn, when the virus is not active," Pernet said.

But warmer waters would reduce this window of opportunity, he said, and new pathogens could arrive if carried north by fish and other sea life fleeing rising temperatures further south.

Adding to the challenges, rising ocean acidity requires oysters to spend more energy in building their shells, Pernet said.